Tower-cap

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Tower caps are a type of mushroom-like, subterranean tree. Once fully grown, they can be designated for wood cutting and produce Tower Cap logs.

[edit] Tree Farms

It is possible for tower caps to spontaneously grow in muddied subterranean areas only if you have discovered an underground river or underground pond. Such features can be located using the site finder and on the local view during embarkation, if the relevant init and world generation parameters are set.

On the subject of spawning Tower Caps, Toady said:

Right now, you can't plant tower-caps or bring spawn along, so you need to find a cave river or cave pool, in which case they will start growing underground if it's all working. It's trickier to find such things, so it's mostly impossible to do. Later, you should be able to plant them. There are no arbitrary vegetation caps now.

The "arbitrary vegetation caps" comment is referring to the old limit of 200 tower caps growing by the cave river in the mud.

To design a tower-cap farm, refer to chambered irrigation, which can be used for a variety of purposes.